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Without having widescreen checked, you are stretching a 1024x768 image to 1680x1050, which is quite a stretch horizontally. With Widescreen checked, you are stretching a 1280x1024 image to 1680x1024, which roughly maintains the proper aspect ratio. The "anamorphic factor" allows you to fine-tune the height if you want to fill the screen.
Note - some video cards don't do a very good job of interpolated stretching, and the background becomes rather blocky, or "pixelated" on high-res monitors. If this happens, setting the display resolution to the Aquarium's native resolution of 1280x768 smooths out the background, but at a slight cost to the clarity of the fish.
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